Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel & Wheeler, 2006: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, Bản mẫu:N°, tr. 1-262 (texte intégralLưu trữ 2013-04-13 tại Wayback Machine).
_tight_crop.jpg/){kind=small}